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Disclosure statement No potential conflict of interest was report ed by the author(s). Funding This was work was supported by The Natu ral Science Foundation of the Xinjiang Uygur Autonomous Region (2021D01A147); Sup ported by Basic Scientific R& D Program of Public Welfare Research Institutions of Xin jiang Uygur Autonomous Region of China (KY2021062); Supported by China Agricul ture Research System of MOF and MARA. Credit authorship contribution statement Shijian Bai : Formal analysis, Investiga tion, Data curation, Writing-original draft, Writing-review& editing, Project administra tion, Funding acquisition. Jinge Hu : Inves tigation, Data curation, Soft, Visualization, Supervision, Writing-review & editing. Ming Zheng : Writing-review& editing. Ji uyun Wu : Investigation. Junshe Cai : Inves tigation. Guang Chen : Field management. Ronghua Zhao : Field management. Jiangfei Meng : Writing-review& editing. Author notes Shijian Bai and Jinge Hu contributed equally to this work. References Araujo, W.J., Dias, P.C., Moraes, G.A.B.K., Celin, E.F., Cunha, R.L., Barros, R.S. and DaMatta, F.M. 2008. Limitations to photosynthesis in coffee leaves from different canopy positions. Plant Physiol Biochem. 46(10):884-890. https://doi: 10.1016/j.plaphy.2008.05.005 Azuma, A., Ban, Y., Sato, A., Kono, A., Shiraishi, M., Yakushiji, H. and Kobayashi, S. 2015. MYB diplo types at the color locus affect the ratios of tri/di-hy droxylated and methylated/non-methylated antho cyanins in grapes berry skin. Tree Genet Genomes, 11-33. https://doi: 10.1007/s11295-015-0855-0 Azuma, A., Yakushiji, H., Koshita, Y. and Kobatashi, S. 2012. Flavonoid biosynthesis-related genes in grape skin are differentially regulated by temperature and light conditions. Planta. 234(4):1067-1080. https://doi: 10.1007/s00425-012-1650-x Carbonneau, A. and Huglin, P. 1982. Adaptation of
training systems to French region. In: ‘University of California, Davis, Grape and Wine centennial Sym posium’ Ed. A.D. Webb (University of California: Berkley, CA) pp.376-385. Carreño, J., Martínez, A., Almela, L., and Fernández López, J.A. 1995. Proposal of an index for the objec tive evaluation of the color of red table grapes. Food Res Int. 28:373-377. Carreño, J., Martínez, A., Almela, L., and Fernández López, J.A. 1996. Measuring the color of table grapes. Color Res Appl. 21:50-54. Castellarin, S.D., Di Gaspero, G., Marconi, R., Nonis, A., Peterlunger, E., Paillard, S., Adam-Blondon, A.F., and Testolin, R. 2006. Color variation in red grapevines ( Vitis vinifera L.): Genomic organisa tion, expression of flavonoid 3′-hydroxylase, flavo noid 3′,5′-hydroxylase genes and related metabolite profiling of red cyanidin-/blue delphinidin-based an thocyanins in berry skin. BMC Genomics. 7(1):12. https://doi: 10.1186/147-2164-7-12 Colova, C., Bordallo, P., Parker, L., Walters, T. and Le ong, S. 2007. Evaluation of yield, fruit quality and photosynthesis of two training-trellis system and canopy management practices for Carlos and noble muscadine grapes in Florida. OENO-One. 41(1):43 49. https://doi.org/10.20870/oeno-one.2007.41.1.856 Esteban, M.A., Villanueva, M.J., and Lissarrague, J.R. 2001. Effect of irrigation on changes in the anthocy anin composition of the skin of cv Tempranillo (Vi tis Vinifera L) grape berries during ripening. J Sci Food and Agr. 81:409-420. https://doi10.1002/1097 0010(200103)81:4<409::aid-jsfa830>3.0.co;2-h Ezzahouani, A. and Williams, L.E. 2003. Trellising, fruit thinning and defoliation have only small effects on the performance of ‘Ruby Seedless’ grape in Mo tocco. J Hortic Sci Biotech. 78(1):79-83. http://dx.doi.org/10.1080/14620316.2003.11511591 Gomez, C., Terrier, N., Torregrosa, L., Vialet, S., Fournier-Level, A., Verries, C., Souquet, J.M., Mazauric, J.P., Klein, M. and Cheynier, V. 2009. Grapevine MATE-type proteins act as vacuolar H+- dependent acylated anthocyanin transporters. Plant Physiol. 150(1):402–415. https://doi:10.1104/99.109135624 Guidelines for the conduct of tests for distinct ness, uniformity and stability-grapevine ( Vitis L.) (UPOV:TG/50/9., Guidelines for the conduct of tests for distinctness, uniformity and stability-Grapevine, NEQ). NY/T 2563-2014. Jackson, D.I. and Lombard, P.B. 1993. Environmental and management practices affecting grape composi tion and wine quality. A review. Am J Enol Viticult. 44:409-430. Jaakola, L., Maatta, K., Pirttila, A. M., Torronen, R.,
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